Dr. Ulmar Grafe received his PhD in behavioural biology from the Department of Neurobiology and Behavior, at Cornell University, USA and his Habilitation from the Department of Animal Ecology and Tropical Biology at Würzburg University, Germany. He has taught at Cornell University, University of Würzburg, and Universiti Brunei Darussalam. His research has taken him to Brunei, Germany, Ivory Coast, Malaysia, Sambia, Zimbabwe, and the USA.


Teaching Area

Animal Physiology, Tropical Ecology, Parasitology, Herpetology, General Biology


Current Research

Current research interests include: the functional ecology of amphibians, the sensory ecology of blood-sucking insects and their frog hosts, the use of multiple signalling modes in anuran communication, and the resource-service mutualism between bats and pitcher plants.


Research Interests

Tropical Ecology, Herpetology, Parasitology, Sensory Physiology



  • Scharmann, M., D.G. Thornham, T.U. Grafe & W. Federle 2013. A novel type of nutritional ant-plant interaction: Ant partners of carnivorous pitcher plants prevent nutrient export by dipteran pitcher infauna. Plos One 8(5) e63556.
  • Grafe, T.U.,D.Preininger, M.Sztatecsny, R.Kasah, J.M.Dehling, S.Proksch & W.Hödl 2012. Multimodal communication in a noisy environment: A case study of the Bornean rock frog Staurois parvus Plos One 7(5) e37965. doi:10.1371/journal.pone.0037965
  • Grafe, T.U., C.R. Schöner, G.Kerth, A. Junaidi & M.G. Schöner 2011. A novel resource-service mutualism between bats and pitcher plants. Biology Letters 7: 436-439. doi: 10.1098/rsbl.2010.1141
  • Keller, A., K.E. Linsenmair, M.O. Rödel & T.U. Grafe 2009. The importance of environmental heterogeneity for species diversity and assemblage structure in Bornean stream frogs. Journal of Animal Ecology 78:350-358 (Highlighted as cover of journal). 
  • Arch, V.S., T.U. Grafe, M. Gridi-Papp & P.M. Narins 2009. Pure ultrasonic communication in an endemic Bornean frog. Plos One 4(4): e5413. doi:10.1371/journal.pone.0005413. (Highlighted in Nature 459:12).